using numerous derived features including the presence of an ossified extensorial arcus (retinaculum extensorii) on the proximodorsal surface of the tarsometatarsus
oldest evidence for daytime behavior in owls
a potential Miocene origin of nonnocturnal habits
demonstrates higher posterior probabilities of diurnal behavior among early diverging Surniini, and the addition of this extinct taxon into analyses enhances the hypothesis of this clade’s diurnal origin
recent phylogenomic analyses support the hypothesis of the independent acquisition of predatory behaviors and diets along with their nocturnal habits
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